The site was originally a British Army and later RAF establishment at Tŷ Croes. Among its earlier history, the camp housed a Royal Artillery training facility and then a TA Barracks. The RAF subsequently operated a radar establishment there, complete with domed arrays and a small narrow-gauge railway. The site was decommissioned around 1992–93.
It was the Wirral 100 motor racing club that first identified the opportunity for motorsport use of the complex's roadways. The first events took place in October 1992 and focused primarily on motorcycle classes. Cars and sprint events followed, and rallycross became a regular activity, including rounds of the British Rallycross Championship. Facilities at this stage were basic but the track surface was noted for its quality.
By 1997 it was clear that racing had a permanent future on Anglesey. Upgrades were carried out to meet full Motor Sports Association (MSA) and ACU (Auto-Cycle Union) licensing requirements, and a small permanent pitlane was installed. The track then became a regular fixture for two- and four-wheeled club racers from Wales and the North West of England.
In 2006 the circuit underwent a far more ambitious overhaul, which saw an almost entirely new course created with four configurable layouts. Only the original start-finish straight and a corner known as Beth's Bend were incorporated into the new design. The four resulting configurations are:
International GP Circuit: 3.380 km (2.100 miles), 11 turns, clockwise
Coastal Circuit: 2.494 km (1.550 miles), 10 turns
National Circuit: 1.931 km (1.200 miles), 6 turns
Club Circuit: 1.287 km (0.800 miles), 4 turns
The International GP configuration runs across undulating ground overlooking the Irish Sea, with Snowdonia visible across the water on the mainland. The layout incorporates cambers, fast sections, technically challenging mid-range corners, hairpin bends, and a banked corner. Racing motorcyclist Peter Williams described it in 2006: "The journey there is breathtaking and the circuit itself is the most scenic in the UK."
In 2019, planning applications were submitted to expand the paddock, garaging, and hospitality facilities at the venue.
While no major national championships have made the circuit a permanent round — its relative remoteness is often cited — Anglesey has hosted a broad range of club car and motorcycle events. The largest annual event is the Racing Rocks weekend, organised by the BRSCC, which culminates in a VW Fun Cup Race into the Night and includes performances from approximately 30 rock groups.
The Race of Remembrance, an endurance race commemorating armed forces personnel, has been held at Anglesey since 2014.
The TV motoring programme Fifth Gear regularly used the Anglesey circuit for its Shoot Out segment, with the Coastal layout in particular favoured for its challenging downhill corkscrew section.
The International GP Circuit lap record stands at 1:21.287, set by Sean Walkinshaw in a Dallara F302 in 2012 during an F3 event. On the Coastal Circuit the outright record is 1:01.741, set by Sylvester Mullins in a Gould GR37 in 2009.
The circuit pays tribute to Tom Pryce, the late Welsh Formula One driver who died during the 1977 South African Grand Prix. One of the straights bears his name. Since 2019 the venue has hosted the Tom Pryce Memorial Trophy, run by the HSCC, commemorating one of Wales's most significant figures in international motorsport.
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