The Tuscan Speed Six was introduced in 1999. The high-performance Tuscan S model was the top-of-the-line variant, rated at 390 hp (395 PS) and featuring aerodynamic improvements including a rear lip spoiler to increase downforce.
A facelift in 2005 resulted in the Tuscan 2, with exterior changes including a redesigned front grille and headlamps, and more conventional taillamps. Mechanical revisions included revised spring rates, improved steering response, and altered suspension geometry to improve on-road drivability. The base models were detuned to 350 hp (355 PS) and 393 N⋅m (290 lb⋅ft) while maintaining a weight of 1,100 kg (2,425 lb). The interior was also refreshed with a more ergonomic layout. Production continued until TVR’s closure in 2006.
Five different inline-six engine options were available to customers. Four of these were naturally aspirated 4.0 L variants of the Speed Six engine, using multipoint fuel injection and producing varying power and torque outputs depending on the trim level. The fifth was a 3.6 L shortened-stroke version of the Speed Six, producing the same power as the lowest-level 4.0 L engine, but with slightly less torque. The engines featured a bore x stroke of 96 mm × 83 mm (3.78 in × 3.27 in) for the 3.6 L version, and 96 mm × 92 mm (3.78 in × 3.62 in) for the 4.0 L versions.
Despite numerous adjustments to the Tuscan’s chassis and suspension, the overall size and appearance of the variants remained largely unchanged, aside from minor aerodynamic aids on the S model, including an undertray and a small boot-lid spoiler. A “Mk 2” version was introduced in October 2005, consisting of cosmetic changes to the front and rear lights, the dashboard, and the spoilers on the S model, alongside minor chassis revisions to improve handling. A full soft top variant was introduced alongside the original targa version at this time.
Testing showed the Tuscan S could achieve 0–30 mph (48 km/h) in 1.72 seconds, 0–60 mph (97 km/h) in 3.68 seconds, 0–100 mph (161 km/h) in 8.08 seconds, and 100–0 mph in 4.15 seconds. These results were achieved in a post-2003 Tuscan S without traction control or anti-lock brakes, features TVR’s design philosophy considered detrimental to performance and safety. TVR rejected the notion that these features, along with airbags, were “safety devices”, believing their cars were safer without them based on testing and experience.
A modified version of the car competed in the 2003 and 2004 editions of the 24 Hours of Le Mans.